I was happy to see recently that Robert Casey has done a study on Z16343. He was primarily looking at YDNA STRs which are better tested but less predictable than SNPs. Z16343 is important for my branch of Hartleys as it is under that branch.
Where is Z16343?
Z16343 is under R-L513, which is under L21 which is under R1b. Here is page 1 of 2 of R-L513:

If the two pages were to be put together, Z16343 would be in about the middle of L513. The circle indicates where Z16343 is. It is a fairly small branch. The names circled represent only the branches tested by the BigY SNP test. Here is a closer look at Z16343:

Here I see only the following 13 surnames:
- Pilsbury
- Smith
- Hays
- Goff
- Merrick
- Hartley
- Hays
- Davis
- Williams
- Martin
- Thomas
- Philips Bennett
- McCullers
These surnames are in 11 Haplogroups. Four Haplogroups contain two surnames:
- BY11565 – Goff and Merrick
- FTB44077 – Davis and Willliams
- FGC33966 – Martin and Thomas
- FT135561 – Bennett and McCullers
There are two likely reasons that there would be more than one surname per haplogroup. One is that the haplogroup formed before the time of surnames. When surnames were adopted, different surnames would have been used for people in the same haplogroup. The other reason is due to a NPE, a non-paternal event. This would be due to an adoption or single mother situation for example where the male surname would have changed over the years.
Due to additional testing, two Surnames have additional branching:
- Hayes – ZS349 and BY13845
- Hartley – A11132 and FT225247
Under Hartley, where I have particular interest, there were four testers. I tested my brother to get a terminal haplogroup named. There are two other Hartleys that are under A11132.
There are three empty haplogroups. These are at branching points:
- Z16343
- Z16854 (the side with Pilsbury and Hays)
- Z17911 (the side with all but Pilsbury and Hays)
- FT9480 (a branch with all but Pilsbury, Hays, Smith and Hartley)
SAPP
Robert Casey uses the SAPP program for his analysis. Here is the SAPP web page:

The program is by David Vance. Here is Casey’s results for Z16343:

This tree is larger than the BigY SNP tree, because it includes more testers. Casey’s report says:
Z16343 has 19 testers that were confirmed to be Z16343 via YSNP testing. Another 59 testers were predicted for a total of 75 testers.
I have not used the SAPP program myself. I notice that it has a SNP Tree function, so I tried that for Z16343 and came what looks to me to be an upside down tree:

I’m used to having the older branches at the top, but this one has the older branch at the bottom. When I add names, I get this:

When I compare this tree to the L513 Tree, I see that Davis and Williams are missing in a parallel branch to Martin and Thomas in the above tree. Also Hays at ZS352 is ZS349 in the L513 Tree. The thing I don’t like about this tree (and other trees) is that it seems to imply that Hartley descends from Smith. In actuality, Hartley and Smith descend from a common ancestor who lived before the time of surnames. According to YFull, A11138 formed 1750 years before present or around the year 272! However, YFull does not have the full results of those who have completed the BigY test.
More on Casey’s SAPP Tree
Casey mentions the signature for Z16343 and I think I can pull it off the tree:

I will ignore the ReclOH as I don’t understand that very well (sorry McCullers). One problem with the SAPP tree is that is seems to imply that the McCullers Branch is directly under Z16343. However, by the SNP tree it is really Z16343 > Z17911 > FT94840 and so on. Jared Smith has recently drawn his own tree which includes the parent SNP to Z16343 which I have copied below:

The first branch after the McCullers Branch has Pilsbury (shown above) and Hays (not shown). I’ll put the STRs from above Node #99 (or BY13850} into a small spreadsheet:

Not counting the ReclOH, there are 7 main branches under Z16343 on the SAPP generated YDNA Tree. Node #150 is one of those 7 major branches, but it has no STR deviations from the signature for Z16343. However, it’s two sub-branches have deviations

I’ll show them like this in my spreadsheet:

So far I have 9 STRs in the Z16343 Signature. Casey said that there were only 8 STRs in the signature:
The signature of Z16343 includes eight markers which is acceptable. But this signature does not include slow mutating markers or multi-step mutations and is not the strongest signature.
Here is my spreadsheet for the seven main branches under Z16343:

This shows 12 STRs in the signature. I added in dates for the branches. Pilsbury has the latest date for BY13850, but also the most variations from the signature Z16343 STR. I highlighted the two 12’s under 439. That could normally be a problem, but these were apparently determined to be parallel mutations as the SNPs showed that they were in a different branch.
The Hartley Branch of the SAPP Tree
As I know most about the Hartleys, I will look at what Casey’s SAPP Tree did for them:

I have started by highlighting my branch which I feel cannot be right. My brother and I have the obviious ancestor of my father. He was born in 1918. The date given is 1900 which is not far off. The 1-3 generations is right. However, it is the relationship with Steve that seems off. Here is my brother Jim’s block tree:

This shows that Jim and I have 7 SNPs up to our common ancestor. Steve has 5 Private SNPs but only 2 average SNPs. If I say there are about 5 SNPs up to our common ancestor and I use 83 years per SNP, then that is 415 years. If I take that time from 1960, then that goes back to the year 1545. Another check is Michael’s genealogy. His ancestor Edmund Hartley was born in 1666 in Lancashire County, England and came to Pennsylvania around the year 1700. His father Roger Hartley was born in 1628. I don’t think that I descend from Roger, but if I did, that would mean if the block tree is right, I couldn’t have a common ancestor with Steve and Michael any later than 1628.
The SAPP Tree and Steve
The SAPP Tree shows a much later date for the common ancestor between me, my brother and Steve. By this I mean much later than seems to be warranted given the YDNA SNPs:

That shows a range between 1750 and 1850. That could be off by 3-400 years. Both the SNPs and our genealogy suggest that 1800 cannot be right.
The SAPP Tree and Michael
However, the SAPP Tree for Michael shows a common ancestor which appearst to be too far away.:

Node #121 shows a date between 1200 and 1550. The SNPs seem to suggest that both Michael and Steve and I should have a common ancestor around Node #101 or around the year 1500.
The Other Three Hartleys on the SAPP Tree

I’m curious as to how the SAPP Tree would have come out with John’s new 111 STR results. Here, he is shown as further from me compared to Steve. John is my closest match other than my brother at the 111 STR level:

The TiP report gives a date to a common ancestor with John at about 1530, so that part is not so far off. It is more that Steve is shown as matching more closely than he should be. The SAPP tree also gives a common ancestor between me and John at the year 1500 which seems reasonable:

Lawrence has recently put in for a BigY 700 test, so he is a very important YDNA tester.
My Previous Attempts at Hartley STR Trees
I have made a few attempts at drawing my own Hartley STR trees. Here is one that I built:

This looks somewhat similar to the SAPP tree. This reminds me of my frustration at having Steve test and finding out that he was not related as closely to me by SNPs compared to what the STRs seemed to show. The common answer to the discrepancy lies in back mutations and parallel mutations which are difficult to detect. In my tree, Michael is on the right and in the SAPP tree, Michael is on the left. However, the tree is basically similar. In the tree above, I tested as well as my brother. Interestingly, I had a new STR mutation in 1956 when I was born at STR 534. Steve has completed his BigY test. Lawrence ordered a BigY recently as did John. However, I don’t see John on my chart. That may be because he tested to 67 STRs and this could be a 111 STR chart.
John is on the SAPP Tree:

The tree shows our common ancestor at Node #101. However, once Steve’s BigY information comes in, it will clear up the tree.
This is the SNP working model I am using:

In this model, with the information I now have, we are about at the same level for a common ancestor. My guess is that our common ancestor is around 1500. Actually, that is true for all but John. John and Lawrence are new testers, but Lawrence’s dating should be tied to Michael’s.
More on John’s STRs
John’s STRs were updated recently as part of his BigY order. I wrote a short Blog on those results here. My best guess using FTNA’s TiP Report is that John and I should have a common ancestor around the date of 1565 (although I also have a 1530 estimate above). The interesting this is, that by STRs John is related most closely to my brother Jim and then me. This is because I had an extra STR mutation that my brother Jim did not have. Then after that, John shows a more distant match with Steve. There are more Hartleys out there that have tested and are related to us by YDNA, but the STR test does not show a match with them because they are beyond the FTDNA matching limit. That includes Michael who has taken the BigY test.
Summary and Conclusions
- In my Blog above, I took a rambling look at Casey’s SAPP tree analysis of Z16343
- I don’t believe that the SAPP Report that Casey did was meant to supersede work already done with trees based on SNPs
- Casey’s main point was that STRs should be used to increase the pool of predicted families under Z16343. Based on his study, he went from 19 testers under BigY testing who would be under Z16343 to 75 who should be under Z16343 by STRs.
- The tree works out overall, but in the details where there are discrepancies between the SAPP STR tree and the SNP tree, the SNP tree should be used.
- Every study and analysis that is done on the exisitng YDNA testing brings a little more understanding and light to the architecture of the Z16343 tree.